2.3.1->2.5.1 fails with "Invalid default value for \'DateExpires\'"

Hi,
Using a test server, I overwrote the 2.3.1 files with the 2.5.1 files, cleared the cache & removed the settingscontroller (as per the 2.5 update README).
When I ran mydomain.com/utility/update, I got a 'something went wrong error', so I enabled DEBUG mode and re-ran it.

Any ideas on what I can do? (I'm assuming that I'd get the same error running this on the main site). The 2.3.1 install was a new install btw.

This is the DEBUG output, which seems to indicate a default value problem in the DB:

Invalid default value for 'DateExpires'
The error occurred on or near: /home/mydomain.com/public_html/library/database/class.database.php

403: // If we get here then the pdo statement prepared properly.

404: break;

405:

406: } catch (Gdn_UserException $uex) {

407: trigger_error($uex->getMessage(), E_USER_ERROR);

408: } catch (Exception $ex) {

409: list($state, $code, $message) = $pDO->errorInfo();

410:

411: // If the error code is consistent with a disconnect, attempt to retry

Backtrace:

/home/mydomain.com/public_html/library/database/class.database.phpPHP::gdn_ErrorHandler();

[/home/mydomain.com/public_html/library/database/class.database.php:407] PHP::trigger_error();

[/home/mydomain.com/public_html/library/database/class.databasestructure.php:442] Gdn_Database->query();

[/home/mydomain.com/public_html/library/database/class.mysqlstructure.php:313] Gdn_DatabaseStructure->executeQuery();

[/home/mydomain.com/public_html/library/database/class.databasestructure.php:509] Gdn_MySQLStructure->_create();

[/home/mydomain.com/public_html/applications/dashboard/settings/structure.php:292] Gdn_DatabaseStructure->set();

[/home/mydomain.com/public_html/applications/dashboard/models/class.updatemodel.php:666] PHP::include();

[/home/mydomain.com/public_html/applications/dashboard/controllers/class.utilitycontroller.php:290] UpdateModel->runStructure();

[/home/mydomain.com/public_html/library/core/class.dispatcher.php:833] UtilityController->update();

[/home/mydomain.com/public_html/library/core/class.dispatcher.php:262] Gdn_Dispatcher->dispatchController();

[/home/mydomain.com/public_html/index.php:29] Gdn_Dispatcher->dispatch();

Variables in local scope:

[sql] 'create table GDN_AccessToken (
AccessTokenID int not null auto_increment,
Token varchar(100) not null,
UserID int not null,
Type varchar(20) not null,
Scope text null,
DateInserted timestamp not null,
InsertUserID int null,
InsertIPAddress varbinary(16) not null,
DateExpires timestamp not null,
Attributes text null,
primary key (AccessTokenID),
unique index UX_AccessToken (Token),
index IX_AccessToken_UserID (UserID),
index IX_AccessToken_Type (Type)
) engine=innodb default character set utf8mb4 collate utf8mb4_unicode_ci;'

[inputParameters] NULL

[options] array (
)

[returnType] NULL

[tries] 2

[try] 0

[pDO] array (
)

[pDOStatement] false

[state] '42000'

[code] 1067

[message] 'Invalid default value for \'DateExpires\''

[uex] array (
)

Additional information for support personnel:

Application: Vanilla
Application Version: 2.5.1
PHP Version: 7.0.28-0ubuntu0.16.04.1
Operating System: Linux
Server Software: Apache
User Agent: Mozilla/5.0 (Macintosh; Intel Mac OS X 10.13; rv:59.0) Gecko/20100101 Firefox/59.0
Request Uri: /utility/update
Controller: PHP
Method: gdn_ErrorHandler